The
Green River Formation comprises several basins formed as part
of the uplifting of the Rocky Mountains during the Lower Teritary.
These outcrops that occur in Wyoming, Colorado and Utah are
a Laggerstatten in the truest sense, with an enormously diverse
faunal and floral assemblage of fossils exhibiting remarkable
preservation.
The
Green River Formation is actually a heterogeneous complex of
lakes differential in ecological, geological characteristics,
timeframe and hence fauna and flora. The complex comprises three
primary lakes formed as a consequnce of drainage from tectonic
highlands envolved in the uplift of the Rocky Mountains during
Tertiary time. Fossil
lake, centered in Southwest Wyoming, is the smallest and appeared
briefly during the early Eocene. The Lake Gosiute
deposits span the period from Lower to Middle Eocene, and the
largest deposit from Lake Uinta that ranges across the Utah-Colorado
border, spans most of the Eocene Epoch.
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The
bat fossil Icaronycteris, shown above, is the oldest known flying mammal in the
fossil record. |
While
the Green River Formation is most known for its exquisite fish fossils, the fossil
assemblage fully represents the entire Tree of Life, including stromatolites,
plants, invertebrates and vertebrates (including reptiles, mammals and even primates).
One of the most famous and exceedingly rare fossils is Icaronycteris, the oldest
known flying mammal fossil. The index fossil shown in the adjacent picture was
preserved with its skeleton, membranes cartilage, ingested food and waste all
in situ.
During
the Eocene, based on the fossil record, the region was sub-tropical
to temporate. Some 60 vertebrate taxa have been described from
the formation, as well as abundant invertebrates and plants.
Green river has been noted for its well-preserved fish since
mid-way through the 19th century. The unusually excellent preservation
of the Green River fish
fossils is usually attributed to a combination of two factors:
1) a cold period during the Eocene that would have caused dead
fish to sink faster due to a less inflated swim bladder; and
2) the great depth of the lakes and the consequent anoxic conditions
that would have often prevented scavengers from disturbing the
carcasses.
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Exceedlying
rare Green River gar fish: Lepisosteus simplex - 29 inches in length. |
The
majority of fish fossils are taken from the Fossil Lake area from two layers:
1) the so-called 18-inch layer; and 2) the spilt fish layer. The best preserved
fish come from the 18-inch layer. Becuase the sediment is highly laminated, the
fish can often be removed nearly whole. This layer, in the area near Fossil Butte,
does indeed avarage about 18 inches in thickness, and represents some 4000 of
deposition. The composition of the limestone indicates that the layer was formed
in deep water far from shore. By contrast, the so-called split-fish layer is unlaminated
making extraction and preparation of the best fossil fish far more difficult.
The layer is about six feet thick, and the fauna indicates water that was better
circulated than that associated with the 18-inch layer. Some 19 genera of Eocene
fossil fish come from the Green River formation.